Infamous vigilante and anti-hero Frank Castle (Ray Stevenson) is six years into his vengeance-driven zeal as the Punisher. When he brutally assaults a "beat the rap" party for notorious mob boss Gaitano Cesare, he is alerted to the fact that Billy Russoti (Dominic West) has escaped to his recycling plant hideout by Detective Saffiotti (Tony Calabretta) who along with Detective Martin Soap (Dash Mihok) were staking out the party. Castle infiltrates Russoti's hideout and murders Nicky Donatelli (Romano Orzari) as he is exiting the bathroom. The ensuing gunshot alerts Russoti, and a brief fire fight between Castle and he ensues, ending in Russoti being thrown into a glass-crushing machine which ends up leaving the gangster hideously disfigured. Castle, who has taken cover behind the body of Donatelli, realizes that the man he killed was an undercover FBI Agent.
Agent Paul Budiansky (Colin Salmon), the ex-partner of the undercover agent, joins the NYPD's "Punisher Task Force" and partners up with Soap to help bring Castle to justice, once and for all. Meanwhile, a facially mutilated Russoti recovers from his run-in with the Punisher with revenge on his mind and frees his deranged brother, Loony Bin Jim (Doug Hutchison) under his new alias "Jigsaw."
Distraught that he has now become the very evil he swore to battle, Castle attempts to make reparations to Donatelli's wife, Angela (Julie Benz) and daughter, Grace (Stephanie Janusauskas) to no avail. Castle considers retiring from the vigilante business. However, upon hearing information from his partner and armorer, Microchip (Wayne Knight), plans quickly change once he learns that Jigsaw has learned of Donnatelli's family's existence, and will be going after them for revenge.
Jigsaw, Loony Bin Jim, Ink, and Pittsy break into Donatelli's house and hold the family hostage while they tear the house apart looking for the money Donatelli was in charge of. Meanwhile the Punisher goes after the 3 criminals led by Maginty, (T. J. Storm) who were hired to run the delivery for Jigsaw's "biological goods". While doing an acrobatic roll through the air, one is shot by a grenade launcher, another is shot by a rifle, and Maginty has his knees shot out and interrogated by the Punisher as to Jigsaw's whereabouts. Maginty informs Jigsaw went after Donatelli's money so he could pay them for the job. Begging for his life, the Punisher doesn't shoot him, but instead throws him off the roof and onto a spiked fence, killing him.
Agent Budiansky arrives just in time with Detective Soap (who learned of the Punisher's whereabouts by going after the first C.B. call they heard about the three criminals). Agent Budiansky attempts to arrest the Punisher but is easily disarmed and the Punisher warns him to leave him alone but just as he is about to walk away Budiansky handcuffs himself to the Punisher. The Punisher points out that Jigsaw went after Donatelli's family and Budiansky has to let him go, but Budiansky just sents one patrol car to check on their house(and the cops are easily killed) and intends to take the Punisher back to the station instead, but is convinced to check on the house himself after the patrol car fails to respond.
Budiansky enters the house, refusing the Punisher's help, and is forced to drop his weapon by the sight Ink and Pittsy holding the family hostage. Soap frees the Punisher, who kills Ink, and Budiansky disarms Pittsy and is about to handcuff him when the Punisher shoots Pittsy in the face with a shotgun, blowing apart the handcuffs in Budiansky's hand in the process. Budiansky can only draw his weapon and point it at the Punisher in protest to no avail. Budiansky then arrests Jigsaw and Loony Bin Jim after a short gun fight in which the 2 criminals simply empty their clips at Budiansky who was behind cover and still had ammunition left.
Jigsaw and his brother escapes custody by giving up Cristu Bulat (David Vadim), who was smuggling in a biological weapon to Arab terrorists in Queens to the FBI, in exchange for total immunity, the money Donatelli hid and a file on Microchip. They take Microchip hostage, killing his mother in the process and hiring a small private army of criminals, and then holed up in the Bradstreet Hotel.
Castle enlists the help of Agent Budiansky, who sets up a distraction for him at the Bradstreet Hotel by offering up Jigsaw to Tiberiu Bulat (Aubert Pallascio) Cristu's father who sends in his goons to start a shootout in the hotel's lobby. Before entering, Castle knocks Budiansky out, knowing that he must face Jigsaw alone. Once inside, a brutal shootout ensues with Jigsaw's goons. Castle confronts Jigsaw and his insane brother, both holding Microchip and the Donatelli family at gunpoint. Jigsaw gives Castle a choice: If Frank shoots one of his friends, Jigsaw will let the others go free. Microchip boldly offers his life to save the Donatellis, but Castle chooses to shoot Loony Bin Jim instead. As a result, Jigsaw kills Microchip. Enraged by the loss of his partner, Castle attacks Jigsaw, ultimately ending with the Punisher impaling Jigsaw with a metal rod and then throwing him onto a fire. As Jigsaw burns to death, Castle calmly tells him that "This is just the beginning."
Outside the building, Castle bids farewell to the Donatelli family. As they are walking down the street, Soap tries to persuade Frank to give up his role as the Punisher. Soap has a change of heart, however, when he is attacked by a mugger, who quickly becomes another victim of the Punisher.
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